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bolet.jpg - 4838 BytesTaxonomic Digest of Tylopilus

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Updated: 11 May 2008

Tylopilus
(including Porphyrellus and some Rubinoboletus)

Pileus dry, glabrous to subtomentose, microscopically a trichodermium or subhymeniform.Context white, unchanging or staining pale brown, red then black, or rarely blue. Hymenophore adnexed, white then pinkish flesh colored to purplish brown to rusty brown, staining brown. Stipe dry, pruinose to glabrous to reticulate, to finely scabrous. Spore deposit pinkish flesh colored to purplish brown, to rusty brown. Spores smooth, fusoid. Hymenial cystida present, usually as pseudocystidia. Clamp connections absent.
Distribution: North Temperate zone, montane Neotropics, northern South America, East Asia, Southeast Asia, Indomalaya, Australia, New Zealand, Africa.
Ectomycorrhiza: Pinaceae, Fagaceae, Betulaceae, Myrtaceae, Casuarinaceae, Myrtaceae, Caesalpiniaceae.

Image links available

  1. acutesquamosus Singer, in Singer, Araujo & Ivory, Beih. Nova Hedwigia 77: 117. 1983
  2. alboater (Schwein.) Murrill, Mycologia 1: 16. 1909
  3. albofarinaceus (W.F. Chiu) F.L. Tai, Syll. Fung. Sinicorum: 757. 1979

  4. alkalixanthus Halling & Amtoft, Brittonia 54(4): 262-265. 2002
  5. alutaceoumbrinus Hongo [as 'alutaceo-umbrinus'], Mem. Shiga Univ. 21: 66 1971
  6. ammiratii Thiers, California Mushrooms: 221-222. 1975
  7. amylosporus A.H. Sm., Non-Gilled Fleshy Fungi: 179. 1973
  8. appalachiensis Singer, Sydowia 2: 38. 1948
  9. arenarius Singer, Persoonia 9(4): 432. 1978
  10. areolatus Hongo, Mem. Fac. Lib. Arts Educ. Shiga Univ., Nat. Sci. 12: 42. 1962
  11. areolatus (Berk.) Manjula, Proc. Indian Acad. Sci., Pl. Sci. 92(2): 86. 1983
  12. argillaceus Hongo, J. Jap. Bot. 60(12): 372. 1985
  13. atratus Both, Bull. Buffalo Soc. Nat. Sci. 36: 215. 1998
  14. atrobrunneus (Lj.N. Vassiljeva) Wolfe, Bibl. Mycol. 69: 82. 1979
  15. atronicotianus Both, Bull. Buffalo Soc. Nat. Sci. 36: 216. 1998
  16. austrofelleus (Boletus austrofelleus Cleland, Trans. R. Soc. S. Austr. 48: 246. 1924; (possibly a Tylopilus or Austroboletus, fide Watling & Gregory, Proc. Roy. Soc. Queensland 100: 21. 1988)
  17. badiceps (Peck) A.H. Sm. & Thiers, Boletes of Michigan: 122-123. 1971
  18. ballouii (Peck) Singer, Amer. Midl. Nat. 37: 104. 1947 =Rubinoboletus ballouii (Peck) Heinem. & Rammeloo, Bull. Jard. Bot. Nat. Belgium 53(1/2): 295. 1983
         Rubinoboletus ballouii var. fuscatus (Corner) Heinem. & Rammeloo
         Rubinoboletus ballouii var. luteoporus Heinem. & Rammeloo
         Rubinoboletus ballouii var. viscidus T.H. Li & Watling, Edinburgh J. Bot. 56(1): 145. 1999
  19. beelii Heinem. & Gooss.-Font., Bull. Jard. Bot. Brux. 21: 324. 1951
  20. brachypus Singer, J. García & L.D. Gómez, Beih. Nova Hedwigia 102: 47. 1991
  21. brevisporus Watling & T.H. Li, nom. prov., Australian Boletes - a Preliminary Survey: 54. 1999
  22. brunneirubens (Corner) Watling & E. Turnbull, Edinburgh J. Bot. 51(3): 332. 1994
  23. brunneus (McNabb) Wolfe, Bibl. Mycol. 69: 86. 1979
  24. bulbosus Halling & G.M. Muell., Harvard Pap. Bot. 6: 109. 2001
  25. cartagoensis Wolfe & Bougher, Aust. Syst. Bot. 6(3): 191. 1993
  26. castaneiceps Hongo, J. Jap. Bot. 60(12): 375. 1985
  27. castanoides Har. Takah., Mycoscience 43:397-403. 2002
  28. cellulosus Singer, Amer. Midl. Nat. 37: 105. 1947
  29. chlorinosmus Wolfe & Bougher, Aust. Syst. Bot. 6(3): 207. 1993
  30. chromapes (Frost) A.H. Sm. & Thiers, Mycologia 60: 948. 1968
  31. chromoreticulatus C.B. Wolfe & Bougher, Aust. Syst. Bot. 6(3): 205. 1993
  32. conicus (Ravenel) Beardslee, Mycologia 26: 253. 1934
         =Mucilopilus conicus (Ravenel) C. B. Wolfe.
         =Fistulinella conica (Ravenel) Pegler & T.W.K. Young
  33. conspicuocystidiata Watling & E. Turnbull, Edinburgh J. Bot. 51(3): 333. 1994
  34. corneri Singer, J. García & L.D. Gómez, Beih. Nova Hedwigia 102: 50. 1991
  35. costaricensis Singer & L.D. Gómez, in Singer, García & Gómez, Beih. Nova Hedwigia 102: 36. 1991
  36. cyaneocinctus (Singer) Grund & K.A. Harrison, Bibl. Mycol. 47: 198. 1976
  37. cyanescens T.H. Li & Watling, Edinburgh J. Bot. 56(1): 146. 1999
  38. cyanogranulifer T.H. Li, Watling & Gregory, in Li & Watling, Edinburgh J. Bot. 56(1): 146. 1999
  39. exiguus T.W. Henkel, Mycologia 91(4): 655-665. 1999
  40. eximius (Peck) Singer, Amer. Midl. Nat. 37: 109. 1947
  41. eximius var. nanus Hongo, Beih. Sydowia 8: 200. 1979
  42. felleus (Bull.) P. Karst., Rev. Mycol. 3: 16. 1881
  43. felleus var. minor (Coker & Beers) Pilát & Dermek, Hribovité Huby: 132. 1974
  44. felleus var. uliginosus A.H. Sm. & Thiers, Boletes of Michigan: 114. 1971
  45. ferrugineus (Frost) Singer, Am. Midl. Nat. 37: 106. 1947
  46.      ferrugineus subsp. vinaceogriseus Snell, E.A. Dick & Hesler, Mycologia 43: 359. 1951
  47. formosus G. Stev., Kew Bull. 15(3): 381. 1962
  48. fumosipes var. glabripes C.B. Wolfe, Bibl. Mycol. 69: 40. 1979
  49. funerarius (Massee) Pegler & T.W.K. Young, Trans. Brit. Mycol. Soc. 76(1): 136. 1981
  50. fuscobrunneus Watling, in Watling & Gregory, Proc. Roy. Soc. Queensland 100: 21. 1989
  51. gomezii Singer, in Singer, García & Gómez, Beih. Nova Hedwigia 102: 30. 1991
  52. griseocarneus Wolfe & Halling, Mycologia 81(3): 342. 1989
  53. guanacastensis Singer, in Singer, Araujo & Ivory, Beih. Nova Hedwigia 77: 110. 1983
  54. hondurensis Singer & M.H. Ivory, in Singer, Araujo & Ivory, Beih. Nova Hedwigia 77: 109. 1983
  55. hongoi Wolfe & Bougher, Aust. Syst. Bot. 6(3): 190. 1993
  56. humilis Thiers, Mycologia 58(6): 815. 1966
  57. indecisus (Peck) Murrill, Mycologia 1: 15. 1909
  58. indecisus var. subpunctipes (Peck) Wolfe, Sydowia 34: 211. 1981
  59. intermedius A.H. Sm. & Thiers, Boletes of Michigan: 109. 1971
  60. isabellescens Murrill, Lloydia 9: 329. 1946
  61. jalapensis Singer & J. García, in Singer, García & Gómez, Beih. Nova Hedwigia 102: 16. 1991
  62. javanicus Henn., Monsunia: 146. 1899
  63. leucomycelinus (Singer & M.H. Ivory) R. Flores & Simonini, Rivista di Micologia 43(2): 121. 2000
  64. lividobrunneus Singer, Beih. Sydowia 7: 103. 1973
  65. louisii Heinem., Bull. Jard. Bot. Brux. 34: 270. 1964
  66. minor Singer, Mycologia 37: 799. 1945
  67. mitissimus Singer & L.D. Gómez, in Singer, García & Gómez, Beih. Nova Hedwigia 102: 17. 1991
  68. montoyae Singer & J. García, in Singer, García & Gómez, Beih. Nova Hedwigia 102: 19. 1991
  69. nanus (Massee) Singer, Agaricales in Modern Taxonomy (3rd ed.): 736. 1975
  70. nebulosus (Peck) C.B. Wolfe, Bibl. Mycol. 69: 42. 1979
  71. neofelleus Hongo, J. Jap. Bot. 42: 154. 1967
  72. neofelleus f. olivaceus Hid. Takah., Trans. Mycol. Soc. Japan 27(1): 95. 1986
  73. nicaraguensis Singer & M.H. Ivory, in Singer, Araujo & Ivory, Beih. Nova Hedwigia 77: 107. 1983
  74. niger (Heinem. & Gooss.-Font.) Wolfe, Bibl. Mycol. 69: 111. 1979
  1. nigerrimus (R. Heim) Hongo & M. Endo, Trans. Mycol. Soc. Japan 17(3-4): 346. 1976
         =Retiboletus nigerrimus (R. Heim) Binder & Bresinsky, Fed. Repert. 113(1-2): 37. 2002
  2. nigricans (Pat. & C.F. Baker) Singer, Amer. Midl. Nat. 37: 106. 1947
  3. nigropurpureus (Corner) Hongo, Mem. Shiga Univ. 23: 40. 1973
  4. nigropurpureus (Corner) Watling, in Watling & Turnbull, Edinburgh J. Bot. 51(3): 336. 1994
  5. niveus G. Stev., Kew Bull. 15(3): 381. 1962
  6. obscureviolaceus Har.Takah., Mycoscience 45: 372-376. 2004
  7. obscurus Halling, Mycotaxon 34(1): 109. 1989
  8. ochraceosquamosus Watling, in Watling & Turnbull, Edinburgh J. Bot. 51(3): 337. 1994
  9. olivaceoporus T.H. Li, Watling & N.M. Gregory in Li & Watling, Edinburgh J. Bot. 56(1): 146. 1999
  10. otsuensis Hongo, Mem. Fac. Lib. Arts Educ. Shiga Univ., Nat. Sci. 16: 60. 1966
  11. pachycephalus (Massee) Singer, in Singer, Araujo & Ivory, Beih. Nova Hedwigia 77: 114. 1983
  12. pacificus Wolfe, Bibl. Mycol. 69: 44. 1979
  13. pakaraimensis T.W. Henkel, Mycotaxon 78: 105-114. 2001
  14. palumanus Wolfe & Bougher, Aust. Syst. Bot. 6(3): 198. 1993
  15. parachromapes T.H. Li & Watling, Edinb. J. Bot. 56(1): 147. 1999
  16. peralbidus (Snell & Beardslee) Murrill, Mycologia 30: 521. 1938
  17. peralbidus f. praeanisatus (Murrill) Singer, Amer. Midl. Nat. 37: 103. 1947
  18. pernanus (Pat. & C.F. Baker) Watling, Micologia 2000: 583. 2001
  19. perplexus Watling, in Watling & Turnbull, Edinburgh J. Bot. 51(3): 338. 1994
  20. pinophilus Wolfe & Bougher, Aust. Syst. Bot. 6(3): 203. 1993
  21. pisciodorus (Murrill) Murrill, Lloydia 11: 29. 1948
  22. plumbeoviolaceoides T.H. Li, B. Song & Y.H. Shen, Mycosystema 21(1). 2002
  23. plumbeoviolaceus (Snell & E.A. Dick) Singer, Amer. Midl. Nat. 37: 93. 1947
  24. porphyrosporus (Fr. & Hok) A.H. Sm. & Thiers, Boletes of Michigan: 98 1971
  25. porphyrosporus var. olivaceobrunneus (Zeller & F.D. Bailey) Wolfe, Bibl. Mycol. 69: 51. 1979
  26. potamogeton Singer, Persoonia 9(4): 433. 1978
  27. potamogeton var. aquarius Singer, in Singer, Araujo & Ivory, Beih. Nova Hedwigia 77: 122. 1983
  28. potamogeton var. irengensis T.W. Henkel, Mycologia 91(4): 655-665. 1999
  29. potamogeton var. mitis Singer, Fieldiana, Bot., n.s. 21: 125. 1989
  30. praeanisatus (Murrill) Murrill, Lloydia 11: 29. 1948
  31. propriorichromapes Wolfe & Bougher, Aust. Syst. Bot. 6(3): 201. 1993
  32. punctatofumosus (W.F. Chiu) F.L. Tai, Syll. fung. sinicorum: 757. 1979
  33. queenslandianus Wolfe & Bougher, Aust. Syst. Bot. 6(3): 200. 1993
  34. rhoadsiae (Murrill) Murrill, Mycologia 36: 122. 1944
  35. rhodoconius (Singer) T.J. Baroni, Both & Bessette, Bull. Buffalo Soc. Nat. Sci. 36: 257. 1998
    rigens Hongo, J. Jap. Bot. 54(10): 303. 1979
  36. roseolus (W.F. Chiu) F.L. Tai, Syll. Fung. Sinicorum: 758. 1979
  37. rubrobrunneus Mazzer & A.H. Sm., Mich. Botanist 6: 58. 1967
  38. rufonigricans T.W. Henkel, Mycologia 91(4): 655-665. 1999
  39. rugulosoreticulatus Hongo, J. Jap. Bot. 54(10): 302. 1979
  40. sanctae-rosae Singer, in Singer, Araujo & Ivory, Beih. Nova Hedwigia 77: 112. 1983
  41. sinicus (W.F. Chiu) F.L. Tai, Syll. Fung. Sinicorum: 758. 1979
  42. snellii Wolfe, Mycotaxon 17: 400. 1983
  43. sordidus (Frost) A.H. Sm. & Thiers, Mycologia 60: 950. 196
  44. sphagnorum (Peck) A.H. Sm. & Thiers, Boletes of Michigan: 123. 1971
  45. spinifer (Pat. & Baker) Watling & Lee, Malayan Nature J. 53(4): 315-322. 1999
         ?=Boletochaete spinifer (Pat. & Baker) Singer
  46. striatulus Heinem., Bull. Jard. Bot. Brux. 21: 320. 1951
  47. suavissimus Heinem. & Gooss.-Font. apud Heinem., Bull. Jard. Bot. Brux. 21: 315. 1951
  48. subcellulosus Singer & J. García, in Singer, García & Gómez, Beih. Nova Hedwigia 102: 32. 1991
  49. subchromapes C.B. Wolfe & Bougher, Aust. Syst. Bot. 6(3): 194. 1993
  50. subfusipes A.H. Sm., Persoonia 7(2): 330. 1973
  51. subniger Singer, J. García & L.D. Gómez, Beih. Nova Hedwigia 102: 38. 1991
  52. subpunctipes (Peck) A.H. Sm. & Thiers, Boletes of Michigan: 119. 1971
  53. subunicolor E.A. Dick & Snell, Mycologia 52(3): 447. 1961
  54. subvinaceipallidus T.H. Li & Watling, Edinburgh J. Bot. 56(1): 147. 1999
  55. tabacinus (Peck) Singer, Mycologia 36: 362. 1944
  56. tabacinus var. amarus Singer, Mycologia 37: 799. 1945
  57. tabacinus var. dubius Singer, Mycologia 37: 799. 1945
  58. tenuis Heinem., Bull. Jard. Bot. Brux. 21: 317. 1951
  59. tenuis var. purpureobadius Heinem., Bull. Jard. Bot. Brux. 21: 318. 1951
  60. umbrosus (G.F. Atk.) A.H. Sm. & Thiers, Mycologia 60: 950. 1968
  61. valens (Corner) Hongo & Nagas., Rep. Tottori Mycol. Inst. 14: 87. 1976
  62. variobrunneus Roody, A.R. Bessette & Bessette, in Bessette, Both, Bessette, Dunaway & Roody, Bull. Buffalo Soc. Nat. Sci. 36: 236. 1998
  63. velatus (Rostr.) F.L. Tai, Syll. Fung. Sinicorum: 758. 1979
  64. veluticeps (Pat. & C.F. Baker) Singer, Amer. Midl. Nat. 37: 105. 1947
  65. venezuelae Singer & Digilio, Lilloa 30: 163. 1960
  66. vinaceipallidus (Corner) T.W. Henkel, Mycologia 91(4): 655-665. 1999
  67. vinaceogriseus Singer, J. García & L.D. Gómez, Beih. Nova Hedwigia 102: 40. 1991
  68. vinosobrunneus Hongo, Beih. Sydowia 8: 198. 1979
  69. violaceus (Beeli) Heinem., Bull. Jard. Bot. Brux. 21: 322. 1951
  70. violatinctus T.J. Baroni & Both, Bull. Buffalo Soc. Nat. Sci. 36: 261. 1998
  71. virens (W.F. Chiu) Hongo, Mem. Fac. Lib. Arts Educ. Shiga Univ., Nat. Sci. 14: 46. 1964
  72. virens (W.F. Chiu) F.L. Tai, Syll. Fung. Sinicorum: 758. 1979
  73. viscidichromapes Watling & T.H. Li, nom. prov., Australian Boletes - a Preliminary Survey: 55. 1999
  74. williamsii Singer & J. García, in Singer, García & Gómez, Beih. Nova Hedwigia 102: 33. 1991
  75. zambianus Watling, in Watling & Turnbull, Edinburgh J. Bot. 51(3): 342. 1994

    Sequestrate taxa:

  76. Gastrotylopilus brunneus T.H. Li & Watling, Edinb. J. Bot. 56(1): 145. 1999

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