Leccinum rugosiceps (Peck) Singer, Mycologia 37: 799. 1945.
Pileus 3–10.5 cm broad, at first convex, with age plano-convex, dry, irregularly pitted to rugulose to wrinkled; disc even, at first dark brown to mustard brown to somalis brown, then dark brown to mustard brown to somalis brown to yellowish brown; margin rugulose, when young dark brown to mustard brown to somalis brown to violet brown, then dark brown to mustard brown to somalis brown to yellowish brown; surface subvelutinous; sterile appendiculate flaps absent; with NH4 no reaction. Flesh 10–12 mm thick, white, staining pink to pale red or pink followed by fuscous; odor mild; taste mild. Hymenophore tubulose, depressed. Tubes 10–15 mm long, sordid golden yellow to wax yellow; pores up to 1 mm wide, yellow, then sordid khaki yellow, brown when bruised. Stipe 7–13 cm long, 7–15 mm wide, equal to subequal to subclavate to clavate, dry; upper half when young pruinose, pale yellow, with age scabrous and ridged with pallid ridges, dark sordid yellow; lower half when young subscabrous ridged, white to pallid, with age subscabrous ridged, grayish yellow to pallid; s on upper half when young cinnamon to pale yellow, ge cinnamon to pale yellow, er half when young dark gray to gray, ge dark gray to gray; base white, staining not present. Stipe interior solid; flesh above when young white, with age white, staining pink; flesh at base when young white, with age white, staining pink. Basal mycelium yellow.
Basidiospores 14.7–18.9 µm long, 4.9–5.6 µm wide, mean Q = 3.07, smooth, fusoid to subfusoid, mostly inamyloid (a few dextrinoid), with KOH light brown melleous. Basidia 28–42 µm long, 9–12 µm wide, clavate, hyaline, 4 -sterigmate. Hymenial cystidia 40–60 µm long, 7–9 µm wide, more common towards edge of tubes, thin walled, hyaline, subfusoid to fusoid to ventricose, encrusting pigment absent. Tube trama boletoid, hyaline to golden yellow, lateral strata elements 3.5–6 µm wide, subgelatinous. Pileipellis hyphae hymeniform and forming a palisade (several cells deep), in KOH yellow ochraceous, inamyloid; elements 7–17.5 µm wide, subelongate to subisodiametric or cellular, smooth, thin walled, not gelatinized. Intercalary cells isodiametric to subisodiametric, 7–17.5 µm long, 7–14 µm wide. Pileus trama interwoven, hyaline, inamyloid, with elements 3.5–14 µm wide, smooth, thin walled. Stipitipellis hyphae vertically oriented, parallel, giving rise to clusters of caulocystidia, clavate to ventricose to lageniform to fusoid, with pale yellow brown contents or hyaline, with incrusting pigment absent. Stipe trama hyphae parallel, cylindric, hyaline, inamyloid. Clamp connections absent.
Mycorrhizal host: Quercus.
As the epithet indicates, the pileus surface is rugose; more so in some collections than in others. The hymenophore is pale yellow and the scabers on the stipe are pale caramel colored. The context of the basidiomata is white but soon changes to pink or pale red when exposed. It also belongs in sect. Luteoscabra. Originally described from the eastern United States, L. rugosiceps occurs under Quercus as far south as southern Colombia. In Costa Rica, it seems to be uncommon but widespread in the montane oak forests.
Desc0006.
Cite this publication as: Halling, R. E., G. M. Mueller, and M. J. Dallwitz. 1998 onwards. Leccinum and Phylloporus in Costa Rica: Descriptions, Illustrations, Identification, and Information Retrieval. Version: 13 Decmber 2006. http://www.nybg.org/bsci/res/hall/deltaindex.html. Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993 onwards, 1998) should also be cited (see References).
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