Macleania loeseneriana Hoerold, Bot. Jahrb. Syst. 42: 302.  1909.  Illustration: Luteyn (1996), Plate 2.  Type.  Ecuador.  Pichincha:  Nanegal Valley, Mar 1900 (fl), Sodiro 92/2c (holotype, B†, photos F neg. 4711 and ACS neg. 165;  lectotype, designated by Luteyn (1996), NY fragment ex B).  Image:  Habit.
    Macleania laurina S. F. Blake, Proc. Biol. Soc. Washington 35: 121.  1922.  Type.  Ecuador.  Carchi:  nr El Angel, ca. 3400 m, 5 Jun 1921 (fl), Popenoe 1340 (holotype, US;  NY fragment ex US, photo ACS neg. 63).
        Coarse terrestrial shrub 2-3 m tall, rarely epiphytic, sometimes lianoid, sometimes arborescent to 7 m tall and 6 cm dbh, often arising from lignotuber;  stem terete, conspicuously striate, glabrous;  twigs subterete, bluntly angled, striate or ribbed, densely puberulent, glabrate.  Leaves thick-coriaceous, elliptic-ovate or oblong, 4-17.5 x 2.5-10.5 cm, base rounded, sometimes slightly subcordate, or sometimes very short-attentuate, apex acute or rarely rounded, glabrous or sparsely to densely short-pilose along nerves beneath, also conspicuously punctate on both surfaces from deciduous, sunken glandular fimbriae or these subpersistent beneath;  pinnately nerved with 3-4 lateral nerves per side (or often appearing plinerved), midrib conspicuously thickened and raised in proximal 2-4(-5) cm then impressed distally above, prominently raised beneath, lateral nerves impressed above and raised beneath, reticulate veinlets obscure but impressed above and slightly raised beneath;  petiole subterete, sometimes flattened above, strongly rugose, narrowly winged, 10-20 mm long, puberulent, glabrate.  Inflorescence axillary, racemose, 10-25(-32)-flowered, subtended by a series of usually persistent, broadly ovate to oblong, rounded, densely short-pilose, glabrate, bracts to 12 x 10 mm, the apices not reflexed, the margins scarious;  rachis congested, subterete, often sharply angled, striate, 3-14(-24) cm long and ca. 6 mm diam., densely short, white pilose;  floral bract caducous or deciduous, not reflexed, oblong-elliptic, apically rounded, 11-17(-25) x (3-)7-8 mm, densely short, white pilose dorsally, ciliate, sometimes marginally glandular-fimbriate;  pedicel subterete, conspicuously striate, 16-28 mm long, short-pilose;  bracteoles nearly basal, rarely medial, caducous, oblong-ovate, obtuse to acute, 4-8 x 1.5-3 mm, short-pilose dorsally, ciliate.  Flowers with calyx 8-10 mm long, short, white pilose;  hypanthium subcylindric, striate, rugose, 3-6 mm long, red;  limb spreading, 5-7 mm long;  lobes broadly triangular, acute, 1.5-2 mm long;  sinuses broadly rounded, sometimes tearing;  corolla fleshy, cylindric, terete, 21-25 mm long and 11-16 mm diam., basally red, apically white, densely short, white pilose, the lobes bistratose, slightly spreading, oblong, acute 2-4 mm long, white;  stamen 10, 12-16 mm long;  filaments distinct, 4-5 mm long, glabrous or marginally pilose, the connectives sometimes alternately shouldered;  anthers 9-14 mm long;  thecae 5-8 mm long, the base incurved;  tubules 2, connate most of length, sometimes alternate ones connate midway, 4-6.5 mm long,  dehiscing by clefts ca. 3 mm long;  style glabrous, about as long as corolla, to 31 mm long.  Berry spherical, to 21 mm diam., weakly puberulent.

        Distribution.  Endemic to Ecuador;  lower montane dry and moist forest, montane wet forest, to subpáramo, at (1800-)2700-3500 m altitude.

        Local names:  chaquilulo or chaqui-lulu (Quichua), gualicón, hualicón.

        Uses:  fruit edible;  wood as fuel.

        Cultivated:  NCSC.