Themistoclesia epiphytica A. C. Smith, J. Arnold Arbor. 24: 470.  1944.  Type.  Colombia.  Nariño:  Páramo del Tábano, between Pasto and El Encano, 3200 m, 11 Jan 1941, Cuatrecasas 11899 (holotype, A, photo NY neg. 9639;  isotype, COL).  Image:  Habit.

       Terrestrial or somewhat scandent epiphytic shrub with pendulous branches;  stem subterete, densely brownish-hispid when young, glabrate.  Leaves coriaceous, sometimes concave, ovate, 1.7-2.8 x 0.8-1.4 cm, base obtuse or rounded, apex acuminate and subulate with a 3-7 mm long mucro, margin narrowly to moderately revolute, hispidulous all over when young, glabrate;  inconspicuously 3-5-plinerved, midrib impressed above and raised beneath, lateral nerves obscure, slightly impressed or raised beneath, reticulate veinlets impressed;  petiole subterete, 1-3 mm long, hispidulous, glabrate.  Inflorescence axillary, short-racemose, 4-7-flowered (rarely reduced to 1), surrounded at the base by a series of oblong, acute, ca. 2 mm long bracts;  rachis 3-5 mm long, hispidulous;  floral bract narrowly oblong, ca. 1.5 mm long, apically hispidulous;  pedicel 6-13 mm long, hispidulous;  bracteoles medial, less than 1.5 mm long, marginally hispidulous.  Flowers with calyx 5-6 mm long, ± densely hispidulous with hairs to 1 mm long, also glandular-fimbriate;  hypanthium obconic, slightly 5-angled, 3.5-4.5 mm long;  limb erect-spreading, ca. 1.5 mm;  lobes apiculate, 0.3-0.7 mm long;  sinuses flat;  corolla carnose, terete, 9-10 mm long and 4-5 mm diam., red, glabrous but with lobes hispidulous, the lobes deltate, ca. 1 mm long;  stamen slightly shorter than corolla;  filaments glabrous, alternately ca. 3 mm and ca. 4 mm long;  anthers 5-5.5 mm long;  thecae 1.5-2 mm long;  tubules two times the thecae, dehiscing through clefts 0.7-1 mm long;  nectariferous disc pulvinate, minutely or conspicuously hispidulous with hairs to 0.5 mm long;  style equalling the corolla.  Berry not seen.

        Distribution.  Colombia and Ecuador;  montane wet forest, elfin forest, subpáramo, and Espeletia páramo, at 1830-3800 m altitude.

        Uses:  Ecuador:  "pollinated by a small greenish, shining hummingbird" (Holm-Nielsen 41715).