Themistoclesia inflata A. C. Smith, Mem. New York Bot. Gard. 8(1): 41.  1952.  Type.  Ecuador.  Chimborazo:  Cañon de Río Chanchan, ca. 5 km N of Huigra, 1525-1985 m, 19-28 May 1945 (fl), Camp E-3363 (holotype, US;  isotypes, BM, F, GH, K, MO, NY, P, U;  photo of NY type NY neg. 10143).

        Epiphytic shrub with elongate, pendulous branches;  stem terete, striate, glabrate;  twigs somewhat flexuous, subterete, bluntly ribbed to striate, short-pilose with white hairs.  Leaves succulent, to 4 mm thick, and distally inflated when fresh, drying coriaceous, ovate-elliptic, 3.5-6 x 1.5-3.2 cm, base rounded-truncate or minutely auriculate, apex gradually narrowing into an acumen ca. 1 cm long, margin thickened and slightly revolute, glabrous above, glandular-pilose beneath with hairs sunken;  5-plinerved from the base, midrib and lateral nerves inconspicuously impressed above, midrib raised beneath, reticulate veinlets impressed above, all nerves obscure beneath;  petiole subterete, thickened, rugulose, 2-2.5 mm long, densely short-pilose with white hairs, glabrate.  Inflorescence with flowers supra-axillary, solitary or few (2 ?) in a fascicle with a very short rachis (no rachis apparent on US type);  floral bract oblong, acute, less than 1 mm long, marginally short-pilose;  pedicel 12-17 mm long, densely short-pilose with white hairs 0.2-0.3 mm long;  bracteoles basal, caducis, obscure, subulate, ca. 0.5 mm long.  Flowers with calyx 5-6 mm long, short, white pilose;  hypanthium obconic, inconspicuously 5-angled, 3-4 mm long;  limb suberect, shorter than hypanthium, glabrate;  lobes minute, broadly deltate, 0.5-1 mm long;  sinuses rounded;  corolla thin, broadly cylindric-urceolate, the throat not much constricted, terete, ca. 7 mm long and ca. 4.5 mm diam. at the middle, glabrous, cream-white, the lobes reflexed, deltate, subacute, ca. 1.5 mm long, thus slightly exposing the stamen tips at anthesis;  stamen 10, slightly unequal, about as long as corolla, ca. 5.5 mm long;  filaments distinct, ca. 1.5-1.8 mm long, dorsally hispidulous distally with hairs ca. 0.5 mm long;  anthers 4.5-5 mm long;  thecae 1.7-2 mm long, obscurely mucronate at base;  tubules longer than thecae and very narrow, ca. 2.4-2.5 mm long, dehiscing through clefts ca. 0.5 mm long;  style equal to corolla, slightly thickened at base, ca. 6.5 mm long;  nectariferous disc annular-pulvinate, glabrous.  Berry spherical, short-pilose (mature not seen).

        Distribution.  Endemic to Ecuador and known only from the type collection from "forested valleys" at 1525-1985 m altitude.  Rare and endangered.